Human levels are controlled from a first person perspective. The game de-emphasizes the role of a heads-up display: it lacks a life bar, aiming reticle, and ammunition readout (the ammo readout and aiming reticule can be turned on and off at the player's will, but will be disabled by default.), further adding to the challenge and encouraging the player to find alternate weapons and techniques.
Interspersed with human adventure are levels in which the player controls Kong himself, traversing Skull Island's unique geography, battling various giant monsters while defending Ann. The Kong levels take place in a third-person view, as the player directs Kong to punch, grab and use objects/corpses as weapons. He can also bite, climb, charge, hurl enemies and even pound his chest to go into fury mode. When Kong is sent into fury mode, the sky becomes tinted with a golden hue and Kong becomes more powerful and less vulnerable to attack. Many of the Kong sequences fulfill the role of boss fights, as the giant ape is able to effectively battle the gigantic creatures that Jack's weapons cannot harm.
The game features an alternate ending where Kong can be saved during the Empire State Building shootout and safely returned to Skull Island. The ending was approved by Peter Jackson. To unlock the alternate ending, players must complete the entire game and then go back and play through various maps and earn a total of 250,000 points. It can also be accessed by using the cheat codes.
Plot
In 1933, film director Carl Denham (Jack Black), has gotten hold of a mysterious map, which reveals the secret location of a large island known as Skull Island, located in the far reaches of the Pacific Ocean. Carl hires playwright Jack Driscoll (Adrien Brody) to write his script and plucks a starving, out-of-work actress Ann Darrow (Naomi Watts) to play the part of leading lady and a tramp steamer called the Venture to take them to the island. The ship, controlled by Captain Englehorn (Thomas Kretschmann) arrives at the island on October 12. Three lifeboats containing the cast, crew and sailors are dispatched to the island. Due to stormy seas and large rocks, the lifeboat containing Jack, Carl, Ann, Hayes (Evan Parke), and Briggs smashes into a chunk of rocks, killing Briggs.
Hayes shoots out a distress signal, causing Englehorn to come looking for them via the ship's plane and drop ammo supplies for them. The group, after fighting off giant crabs, head onto a rocky outcrop. Carl suggests shooting some test shots for his movie, asking Ann to scream. Her classic damsel-in-distress style wailing is answered by a loud roar. The party progresses forward, meeting up with the second lifeboat containing Preston (Colin Hanks), Jimmy (Jamie Bell) and Lumpy (Andy Serkis), although it cannot land because of the strong current of the sea. The team continue traversing the island, battling with many vicious creatures, and are eventually forced to split up. After a huge battle with Megapedes and Scorpiopedes through a seemingly abandoned village, Jack and Ann are captured by the island natives.
Jack is tied to a stake, and watches helplessly as Ann is taken by Kong, a 25-foot (7.6 m) gorilla during a native sacrifice. Carl saves Jack and the two give chase. During the dangerous journey through the jungle, they have an encounter with Venatosaurus who are feasting on a dead Ferrucutus. Jack and Carl reunite with Hayes after defeating a pack of Venatosaurus with a .45 caliber Thompson submachine gun. Soon after, they find Preston, Lumpy, Jimmy and Baxter, who are crossing a bridge, but they are attacked by a Vastatosaurus rex. Lumpy is torn apart, Jimmy and Baxter fall down into the crevasse, but Preston gets to the other side. Jack is separated from Carl and Hayes, who tell him to continue looking for Ann. Jack eventually finds Ann, but she is kidnapped by a Terapusmordax. Kong comes to the rescue and saves Ann. Jack continues on into the canyon, where he sees a migrating herd of Brontosaurus, and also battles Megapedes and Scorpio-Pedes. A V. rex attacks the sauropods and Jack meets up with Carl and Hayes, and continue on their path. In the jungle, they save Jimmy, who is being attacked by the Venatosaurs. They eventually get on a raft, where Jimmy tells the group that everyone else is dead. After escaping the Skull Islanders, the team are pursued by two V. rexes. Kong battles and kills them. As the team continue their journey, they enter a swamp, and fight against Udusaurs. After leaving the swamp, Kong interrupts their log crossing and tips them into a huge ravine. Carl's camera is broken and he gives up, heading downstream towards the Venture. Jack, Jimmy and Hayes continue their pursuit of Ann.
Jack eventually saves Ann from a V. rex, and the party attempt to find a long stretch of water of which Englehorn's seaplane can land on. After fighting off some Venatosaurus and a juvenile V. rex in a cave, and leaving a swamp, they finally come across a long stretch of water. Englehorn lands on the water, but is forced to take flight as a V. rex arrives on the scene and chases the group. Ann signals for Kong to come. Eventually, Kong comes to the rescue while Jack shoots some Terapusmordax to distract the carnivore. As Hayes tries to stop the fight, opening fire on Kong, the V. rex charges at Kong and inadvertently steps on Hayes, injuring him. Jack and Jimmy stand over Hayes, who tells Jimmy before he dies to get back to the ship. Jack and Jimmy fight many raptors and head back to the stretch of water and find the seaplane. Jimmy leaves with Englehorn and Jack climbs up into the mountains to save Ann.